Ladytron formed in mid-1999. Keyboardists/programmers Daniel Hunt and Reuben Wu settled in Liverpool after a spate of traveling and DJ work in Japan. They began work on their first single, "He Took Her to a Movie," which was recorded for 50 pounds. With vocals by guest Lisa Eriksson, the release brought a wave of critical acclaim, which the band took advantage of with a tour throughout the continent before beginning work on their debut EP, 2000's Commodore Rock. They met Bulgarian Mira Aroyo that summer, who became their vocalist, while Helen Marnie, who sings and plays keyboards for the group, also joined them. The studio full-length 604 followed a year later, and garnered many positive reviews. 2002 saw the release of the more polished Light & Magic and the mix album Softcore Jukebox. The group returned in the summer of 2005 with the Sugar single, which heralded the rock edge of Ladytron's third album, Witching Hour. Extended Play, which featured remixes of Witching Hour's singles, arrived the following spring. Ladytron moved to Nettwerk for their fourth album, the harder-edged Velocifero, which they produced themselves. Their fifth album, 2011's Gravity the Seducer, returned them to their synth pop roots.